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Recently obtained molecular data from collagen  and mitochondrial DNA sequences  fall in line with the diphyly convergent evolution hypothesis, but have overturned some of the other conclusions obtained from morphology.
These investigations consistently place two-toed sloths close to mylodontids and three-toed sloths within Megatherioidea, close to Megalonyx , megatheriids and nothrotheriids.
They make the previously recognized family Megalonychidae polyphyletic , with both two-toed sloths and Greater Antilles sloths being moved away from Megalonyx.
Greater Antilles sloths are now placed in a separate, basal branch of the sloth evolutionary tree. The following sloth family phylogenetic tree is based on collagen and mitochondrial DNA sequence data see Fig.
Bradypodidae three-toed sloths. Choloepodidae two-toed sloths. The marine sloths of South America's Pacific coast became extinct at the end of the Pliocene following the closing of the Central American Seaway ; this caused a cooling trend in the coastal waters which killed off much of the area's seagrass and which would have also made thermoregulation difficult for the sloths, with their slow metabolism.
Ground sloths disappeared from both North and South America shortly after the appearance of humans about 11, years ago. Evidence suggests human hunting contributed to the extinction of the American megafauna.
Ground sloth remains found in both North and South America indicate that they were killed, cooked, and eaten by humans.
Megalocnus and some other Caribbean sloths survived until about 5, years ago, long after ground sloths had died out on the mainland, but then went extinct when humans finally colonized the Greater Antilles.
Two-toed sloths are slightly larger. Sloths are unusual among mammals in not having seven cervical vertebrae. Two-toed sloths have five to seven, while three-toed sloths have eight or nine.
The other mammals not having seven are the manatees , with six. Sloths have colour vision, but have poor visual acuity.
They also have poor hearing. Thus, they rely on their sense of smell and touch to find food. The outer hairs of sloth fur grow in a direction opposite from that of other mammals.
In most mammals, hairs grow toward the extremities , but because sloths spend so much time with their limbs above their bodies, their hairs grow away from the extremities to provide protection from the elements while they hang upside down.
In most conditions, the fur hosts symbiotic algae, which provide camouflage  from predatory jaguars , ocelots ,  and harpy eagles.
These include biting and blood-sucking flies such as mosquitoes and sandflies , triatomine bugs , lice , ticks and mites.
Sloths have a highly specific community of commensal beetles , mites and moths. Incidentally, it appears that sloths benefit from their relationship with moths because the moths are responsible for fertilizing algae on the sloth, which provides them with nutrients.
Their limbs are adapted for hanging and grasping, not for supporting their weight. Muscles make up only 25 to 30 percent of their total body weight.
Most other mammals have a muscle mass that makes up 40 to 45 percent of the total body weight. On three-toed sloths, the arms are 50 percent longer than the legs.
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The limbs are adapted for suspending the body rather than supporting it. As a result, sloths are completely helpless on the ground unless there is something to grasp.
Even then, they are able only to drag themselves along with their claws. They are surprisingly good swimmers. Generally nocturnal, sloths are solitary and are aggressive toward others of the same sex.
Sloths have large multichambered stomachs and an ability to tolerate strong chemicals from the foliage they eat. The leafy food is digested slowly; a fermenting meal may take up to a week to process.
Sloths descend to the ground at approximately six-day intervals to urinate and defecate see Sidebar: A moving habitat.
Princesa is an inquisitive two-toed sloth, also being looked after at the AIUNAU sanctuary in Colombia. They have a multi-compartment stomach that allows them to effectively digest the tough cellulose that they eat.
This unique feature makes sloths stand out from other mammals whose bone structures do not allow this flexibility. They can also hold their breath for an impressive 40 minutes, suppressing their metabolism to make their heart rate a third of its normal speed.
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